Yup. All dead
Time to start all over again with the loss of two hives. This one I believe was doomed in the Fall. It wasn’t the cold that killed them, per se but it looks like they lost their Queen in the Fall and just didn’t know what to do to stay warm. If their Queen was living, they would have all huddled around her and rotated through a ball cluster. The fact that there are no signs of eggs, larvae or disease makes me think this is the way the end was inevitable.